CLARINA SYRIACA (Lederer, 1855)

GB: Levant Grape Hawkmoth

Deilephila syriaca Lederer, 1855, Verh. (Abh.) zool.-bot. Ver. Wien 5: 195.

Type locality: Beirut, Syria [Lebanon].


BIOGEOGRAPHICAL AFFILIATION

Holarctic; western Palaearctic region. Pleistocene refuge: Monocentric -- Syrian refuge.


ADULT DESCRIPTION AND VARIATION

Male Clarina syriaca, Beirut, Lebanon. Male Clarina syriaca, Lebanon. Photo: © Robert Young, Sphingidae Taxonomic Inventory, NHMUK.

Wingspan: 50--65mm. Smaller than Clarina kotschyi (Kollar, [1849]), resembling Dendrolimus pini (Linnaeus, 1758) (European Pine Lappet). There is considerable variation within the species, especially in ground colour, which ranges from deep reddish brown to pale grey. The transverse lines on the wings are sometimes very faint; distal margin of forewing sometimes without indentations. There is a noticeable sexual dimorphism, with males being lighter in colour and smaller than females.

This species appears to have evolved in isolation in the Levant and spread north to Turkey since the last ice age, where it has come into contact and interbred with the closely related Clarina kotschyi, which spread westwards. It is sometimes difficult to assign individuals from the central and southern regions of Turkey to either C. syriaca or C. kotschyi.


Female Clarina syriaca, Yesiloz, near Alanya, Turkey. Photo: © Tony Pittaway. Female Clarina syriaca, Yesiloz, near Alanya, Turkey. Photo: © Tony Pittaway.

ADULT BIOLOGY

Favours the raised edges of cultivated valley floors where boulder-strewn streams bordered by vine-covered trees and shrubs occur; large numbers may be present under such conditions. It is also found on hillsides and in mountain valleys with shrubs and isolated trees, and in hillside vineyards where, in Lebanon, it occurs up to 1000m (Zerny, 1933; Ellison & Wiltshire, 1939).

In southern Turkey it is common in the valleys of the foothills of the Toros Mountains where they rise from the coastal plain; however, it is absent from the plain itself. Villas and farmhouses with pergolas covered with grapevines are especially favoured, especially where they are set into the edges of rocky slopes.


FLIGHT-TIME

Bivoltine; from May to early July, depending on latitude, and again in August/September. It was common east and north of Alanya, Turkey, during June 2004.


EARLY STAGES

OVUM: Oval, dorso-ventrally flattened, pale greenish yellow. Laid singly on the upperside of vine leaves, often intermixed with those of Theretra alecto (Linnaeus, 1758); however, they are smaller than the latter.


Egg of Clarina syriaca, Yesiloz, near Alanya, Turkey. Photo: © Tony Pittaway.

LARVA: Full-fed 55--65mm. Very similar to that of Clarina kotschyi both in colour and behaviour; retains its basic pattern and coloration throughout its development, but thoracic segments 1 and 2 are retractile in the penultimate instar. However, in newly emerged larvae the horn can be either dark grey or pinkish-orange. Found commonly in suitable areas during June and July, and again in September and October.


Fourth-instar larva of Clarina syriaca, Yesiloz, near Alanya, Turkey. Photo: © Mark Boddington. Full-grown larva of Clarina syriaca, Yesiloz, near Alanya, Turkey. Photo: © Tony Pittaway. Full-grown larva of Clarina syriaca, Yesiloz, near Alanya, Turkey. Photo: © Tony Pittaway.

Hostplants. Species of Vitis and Parthenocissus. However, under natural conditions this species generally favours vines of Vitis vinifera hanging down walls and cliffs, and from trees.

PUPA: 30--36mm. Identical in shape and patterning to that of Clarina kotschyi, but more heavily pigmented.


Pupa of Clarina syriaca, Yesiloz, near Alanya, Turkey. Photo: © Tony Pittaway. Pupa of Clarina syriaca, Yesiloz, near Alanya, Turkey. Photo: © Tony Pittaway.

PARASITOIDS

Eupelmidae: Anastatus bifasciatus (Geoffroy); Tachinidae: Compsilura concinnata (Meigen), Drino (Palexorista) imberbis (Wiedemann).


DISTRIBUTION

From northern Turkey south to northern and western Syria, Lebanon, northern Jordan (Müller et al., 2005a) and Israel (Eisenstein, 1984; Müller et al., 2005b). Also central Cyprus, with a single specimen captured in 1950 at Platres (Ian Kitching, pers. comm 2005).

In southern Turkey this species appears to have spread farther west in recent years and can now be found up to Antalya.

Extra-limital range. None.


OTHER SUBSPECIES

None. The closely related Clarina kotschyi is found in eastern Turkey and Iran.


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