SPHINX YUNNANA Brechlin, 2015 -- Yunnan Pine hawkmoth

Male Sphinx yunnana, 20km N Baoshan village, Wumeng Shan, Yunnan, China, 2200m, vii. 2004 [HOLOTYPE]. Photo: © Ronald Brechlin. Female Sphinx yunnana, 20km N Baoshan village, Wumeng Shan, Yunnan, China, 2200m, vii. 2004 [PARATYPE]. Photo: © Ronald Brechlin. Male Sphinx yunnana, Huili, Sichuan, China, 22.vii.1974. Photo: © Tony Pittaway, IZAS.

TAXONOMY

Sphinx yunnana Brechlin, 2015, Entomo-Satsphingia 8(1): 16. Type locality: China, Yunnan, Wumeng Shan, 20km N Baoshan village, 2200m, vii. 2004, 26°28'N 104°27'E.

Note. Mell (1922a) described brown specimens of what he took to be Sphinx caligineus from northern Yunnan as Sphinx caligineus brunnescens, a subspecies that he claimed was confined to montane forests. Later, however, he also referred specimens from northern Guangdong and southern Hunan to this subspecies (Mell, 1922b) and stated that Sphinx caligineus brunnescens was intermediate ('ein Mittelglied') between the dark brown Sphinx caligineus caligineus (Butler) from Japan and the paler grey Sphinx caligineus sinicus (Rothschild & Jordan) from 'Zocé' (= Sheshan), Zhejiang. On 30.vi.1995, the first author captured a specimen of Sphinx caligineus at Hangzhou (Zhejiang) that was transitional in colour between Sphinx caligineus sinicus and Sphinx caligineus brunnescens. In addition, there is a brown female in the IZAS from nearby Sheshan (Zhejiang) (the type locality of Sphinx caligineus sinicus) that matches the description of Sphinx caligineus brunnescens, while grey specimens of what appear to be Sphinx caligineus have been captured at 2250m in northern Yunnan (Pittaway & Kitching, 2000). We therefore considered it likely that Sphinx caligineus brunnescens was simply a colour form of Sphinx caligineus sinicus, analogous to the brown forms that exist in the European species, Sphinx pinastri Linnaeus (Pittaway & Kitching, 2000).

However, with the description of Sphinx yunnana from Yunnan, the situation with regard to the nomenclature and distribution of this species, Sphinx caligineus brunnescens and Sphinx caligineus sinicus has become even more confusing. The grey specimens of Sphinx caligineus mentioned above, which were captured at 2250m in northern Yunnan, have been confirmed as being Sphinx yunnana. In fact, all individuals of Sphinx caligineus sinicus from Yunnan and Sichuan mentioned in Pittaway & Kitching (2000) are Sphinx yunnana. As no individuals of Sphinx caligineus have ever been confirmed from Yunnan and Sichuan, the brown individuals that Mell (1922a) described from northern Yunnan could well be the brown form of Sphinx yunnana.

No type material of Sphinx caligineus brunnescens could be found in CMNH or MNHU; specimens labelled 'type' in CMNH or 'paratype' in MNHU do not have appropriate data. Danner, Eitschberger & Surholt, 1998, Herbipoliana 4(1): 58, erronously gave the type-locality as 'Bergwälder im Norden der provinz Kuangtung'. The above confusion cannot be fully resolved until the types of Sphinx caligineus brunnescens have been examined.


ADULT DESCRIPTION AND VARIATION

Wingspan: 72--78mm. Larger than the more northern Sphinx caligineus sinicus (Rothschild & Jordan, 1903); also paler and more grey than that species; basal area of forewing upperside distinctly greyer than the rest of the wing, and edged with dark grey (Brechlin, 2015). However, there may also be a brown form in this species, as can be found in the Sphinx pinastri/S. morio group (see above).

Male genitalia unlike those of Sphinx oberthueri (Rothschild & Jordan, 1903) and more similar to that found in Sphinx caligineus sinicus (Rothschild & Jordan, 1903), indicating that this species is probably derived from a relict population of Sphinx caligineus (Butler, 1877).


Male genitalia of Sphinx yunnana, 20km N Baoshan village, Wumeng Shan, Yunnan, China, 2200m, vii. 2004. Photo: © Ronald Brechlin.

ADULT BIOLOGY

Montane areas with a good density of pines (Pinus species).


FLIGHT-TIME

China: 23-27.iv (120km N Dali); 30.vi (Yanyuan); 3-25.vii (Huili; Wumeng Shan); 6-10.vii (Yongsheng); 4.viii (Xichang).


EARLY STAGES

OVUM: Unknown.

LARVA: Unknown.

PUPA: Unknown

Larval hostplants. Probably species of Pinus.


PARASITOIDS

Unknown.


LOCAL DISTRIBUTION

China: Sichuan (Leibo; Huili; Xichang; Yanyuan, 1200m); Yunnan (Yongsheng, 2250m; 120km N Dali; 5km N Hutiaoxia, 2200m).


GLOBAL DISTRIBUTION

Besides Sichuan and Yunnan Provinces, China, also northern Thailand (Inoue et al, [1996]).


Global distribution of Sphinx yunnana. Map: © NHMUK.

BIOGEOGRAPHICAL AFFILIATION

Holarctic; eastern Palaearctic region. Pleistocene refuge: Monocentric -- Yunnan refugium.



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© A.R. Pittaway & I.J. Kitching (The Natural History Museum, London)