ACTIAS Leach, 1815

Actias Leach, 1815, in Leach & Nodder, Zool. Miscell. 2: 25.

Type species: Actias luna (Linnaeus, 1758).

A Holarctic genus comprising about 18 species, of which two or three occur in North America and the rest in the eastern Palaearctic and Oriental regions.

HOSTPLANT FAMILIES: Many trees and shrubs.


ACTIAS SELENE (Hübner, [1810]) -- Indian Moon Moth

UK: Indian Moon Moth; Indian Luna Moth, F: Papillon Lune, D: Indischer Mondspinner, RUS: Pavlinoglazka selena, S: Långsvansad påfågelsspinnare;, NL: Indiase maanvlinder; Maanvlinder, H: Indiai holdasszövő, E: Luna de los actias; Mariposa luna de la India, I: Piccolo pavone di notte; Saturnia minore, N: Indisk månespinner.

Actias selene (Hübner, [1810]), Samml. exot. Schmett. 1: 3, fig. 1.

(Taxonomic note. Many of the subspecies assigned to this species in the past have now been given specific status [See notes on distribution for details]. However, whether Actias ningpoana Felder & Felder, 1862 is a good species or merely an ice-age refugial entity/sibling species which took refuge in southeast China, is still open to question. Post glaciation these two 'taxa' appear to have come together again across a broad zone in southern China, with Actias selene having taken refuge in northern India/south east Asia. Such post-glacial zones of secondary contact and hybridization/merger between once isolated sister species are known as suture zones, and have been reported for a variety of taxa in many biomes (Barrowclough et al., 2019; Ipekdal et al., 2020; Moritz et al., 2009; Portnoy & Gold, 2012).)


Global distribution of Actias selene. Map: © Tony Pittaway.

ACTIAS SELENE EBERTI Rougeot, 1969

Actias selene eberti Rougeot, 1969, Entomops 2(13): 165.

Type locality: Safed Koh, Sud, Kotkai, eastern Afghanistan.


BIOGEOGRAPHICAL AFFILIATION

Holarctic; eastern Palaearctic region. Pleistocene refuge: Monocentric -- Sindian refuge.


ADULT DESCRIPTION AND VARIATION

Wingspan 116--122mm. Very similar to the nominate taxon, of which it may only be a from, but wings more blue-green with very reduced chocolate and cream eyespots. Body white, banded horizontally with purple-brown in line with the costa, which is of a similar colour. The 60mm hindwing tails are yellowish, with some pink. Males have more falcate wings than females.


Female Actias selene selene, India. Photo: © Tony Pittaway.

ADULT BIOLOGY

This nocturnal species inhabits open landscapes with scattered trees and shrubs up to 2400m altitude.


FLIGHT-TIME

Probably univoltine, with adults on the wing from mid to late June (Rougeot, 1969).


EARLY STAGES

Unknown, but probably the same as the nominate subspecies (see Gardiner, 1982 & below images).


Eggs of Actias selene selene, India. Photo: © Tony Pittaway.
Third instar larva Actias selene selene, Kunming, Yunnan, China, 22.viii.2005. Photo: © Tony Pittaway. Final instar larva Actias selene selene, Kunming, Yunnan, China, 22.viii.2005. Photo: © Tony Pittaway.

Hostplants. The nominate subspecies feeds on many species of tree and shrubs, and so may subsp. eberti.


PARASITOIDS

Unknown.


DISTRIBUTION

An eastern Palaearctic species which just penetrates the region in eastern Afghanistan, having been found in the Safed Koh mountain range, Nangarhar Province.

Extra-limital range. None.


OTHER SUBSPECIES

Only three -- selene (Hübner, [1810]), brevijuxta (Nässig & Treadaway, 1997), and taprobanis U. Paukstadt & L. H. Paukstadt, 1999. These are distributed from Pakistan east to eastern China and Taiwan, and south to the Philippines, Borneo and Sri Lanka.

The situation in China is a little confusing. Although this warmth-loving species is confirmed from the south of the country, it is not known for sure just how far northeast it extends due to confusion with other similar-looking green species. Populations north of Zhejiang Province are probably the Chinese Moon Moth Actias ningpoana Felder & Felder, 1862 [still regarded by many as a subspecies of Actias selene]. Those in Heilongjiang Provice, Korea and the Russian Far East may be Actias dulcinea (Butler, 1881).

The following, formally regarded as subspecies of Actias selene (or Actias artemis (Bremer & Grey, [1851])), are now usually treated as distinct species: Actias ningpoana Felder & Felder, 1862; Actias rhodopneuma (Röber, 1925); Actias laotiana Testout, 1936; Actias felicis (Oberthür, 1896); Actias sinensis (Walker, 1855); Actias distincta (Niepelt, 1932); Actias gnoma (Butler, 1877); Actias chapae Mell, 1950; Actias dubernardi (Oberthür, 1897); Actias appollo Röber, 1923; Actias dulcinea (Butler, 1881); Actias neidhoeferi Ong & Yu, 1968; Actias xenia Jordan, 1911; and Actias aliena (Butler, 1879).

A recent analysis of this group in the Russian Far East is given by Zolotuhin (2011).



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